The Calvert Formation, deposited in a shallow sea during the late Oligocene and early to middle Miocene (15-27 million years ago), contains a very rich fossil microflora, both in terms of number of specimens and number of species. Most abundant are pollen of oak, pine, and hickory, but exotic taxa (those that no longer occur in Delaware) are present in all samples of this formation. They include pollen of Engelhardia type, Manilkara, Planera (water elm), Alangium(?), and palms. All of these exotics are genera of subtropical or tropical regions, some occurring now in Central America, Florida, and east Asia. The climate during the deposition of the Calvert Formation was probably subtropical and moist.
The microflora of the Bethany formation and the lower part of the Beaverdam Formation is characterized by a Quercus-Carya assemblage, very few non-arboreal pollen, and Pterocarya and Sciadopitys as exotic constituents. This assemblage has much in common with that of the Brandywine Formation of Maryland and the Eastover Formation of Virginia which are of late Miocene or early Pliocene age. The environment of deposition of the Bethany was probably deltaic, and that of the lower Beaverdam fluviatile.